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This is the second part of a three part series of papers. In this paper, we consider a general class of $N\times N$ random band matrices $H=(H_{ij})$ whose entries are centered random variables, independent up to a symmetry constraint. We assume that the variances $\mathbb E |H_{ij}|^2$ form a band matrix with typical band width $1\ll W\ll N$. We consider the generalized resolvent of $H$ defined as $G(Z):=(H - Z)^{-1}$, where $Z$ is a deterministic diagonal matrix such that $Z_{ij}=\left(z\mathds{1}_{1\le i \le W}+\wt z\mathds{1}_{ i > W} \right) \delta_{ij}$, with two distinct spectral parameters $z\in \mathbb C_+:=\{z\in \mathbb C:\im z>0\}$ and $\wt z\in \mathbb C_+\cup \mathbb R$. In this paper, we prove a sharp bound for the local law of the generalized resolvent $G$ for $W\gg N^{3/4}$. This bound is a key input for the proof of delocalization and bulk universality of random band matrices in [2]. Our proof depends on a fluctuations averaging bound on certain averages of polynomials in the resolvent entries, which will be proved in [10].

We consider a general class of symmetric or Hermitian random band matrices $H=(h_{xy})_{x,y \in \llbracket 1,N\rrbracket^d}$ in any dimension $d\ge 1$, where the entries are independent, centered random variables with variances $s_{xy}=\mathbb E|h_{xy}|^2$. We assume that $s_{xy}$ vanishes if $|x-y|$ exceeds the band width $W$, and we are interested in the mesoscopic scale with $1\ll W\ll N$. Define the {\it{generalized resolvent}} of $H$ as $G(H,Z):=(H - Z)^{-1}$, where $Z$ is a deterministic diagonal matrix with entries $Z_{xx}\in \mathbb C_+$ for all $x$. Then we establish a precise high-probability bound on certain averages of polynomials of the resolvent entries. As an application of this fluctuation averaging result, we give a self-contained proof for the delocalization of random band matrices in dimensions $d\ge 2$. More precisely, for any fixed $d\ge 2$, we prove that the bulk eigenvectors of $H$ are delocalized in certain averaged sense if $N\le W^{1+\frac{d}{2}}$. This improves the corresponding results in \cite{HeMa2018} under the assumption $N\ll W^{1+\frac{d}{d+1}}$, and in \cite{ErdKno2013,ErdKno2011} under the assumption $N\ll W^{1+\frac{d}{6}}$. For 1D random band matrices, our fluctuation averaging result was used in \cite{PartII,PartI} to prove the delocalization conjecture and bulk universality for random band matrices with $N\ll W^{4/3}$.

The goal of this paper is to prove rigorous results for the behavior of genealogies in a one-dimensional long range biased voter model introduced by Hallatschek and Nelson [Theor. Pop. Biol. 73 (2008) 158–170]. The first step, which is easily accomplished using results of Mueller and Tribe [Probab. Theory Related Fields 102 (1995) 519–545], is to show that when space and time are rescaled correctly, our biased voter model converges to a Wright–Fisher SPDE. A simple extension of a result of Durrett and Restrepo [Ann. Appl. Probab. 18 (2008) 334–358] then shows that the dual branching coalescing random walk converges to a branching Brownian motion in which particles coalesce after an exponentially distributed amount of intersection local time. Brunet et al. [Phys. Rev. E (3) 76 (2007) 041104, 20] have conjectured that genealogies in models of this type are described by the Bolthausen–Sznitman coalescent, see [Proc. Natl. Acad. Sci. USA 110 (2013) 437–442]. However, in the model we study there are no simultaneous coalescences. Our third and most significant result concerns “tracer dynamics” in which some of the initial particles in the biased voter model are labeled. We show that the joint distribution of the labeled and unlabeled particles converges to the solution of a system of stochastic partial differential equations. A new duality equation that generalizes the one Shiga [In Stochastic Processes in Physics and Engineering (Bielefeld, 1986) (1988) 345–355 Reidel] developed for the Wright–Fisher SPDE is the key to the proof of that result.

A new non-conservative stochastic reaction–diffusion system in which two families of random walks in two adjacent domains interact near the interface is introduced and studied in this paper. Such a system can be used to model the transport of positive and negative charges in a solar cell or the population dynamics of two segregated species under competition. We show that in the macroscopic limit, the particle densities converge to the solution of a coupled nonlinear heat equations. For this, we first prove that propagation of chaos holds by establishing the uniqueness of a new BBGKY hierarchy. A local central limit theorem for reflected diffusions in bounded Lipschitz domains is also established as a crucial tool.